CFeS 2025(2): 35-39 Bibliography

Published online in:

Chronicles
Fluturat e Shqipërisë
CFeS 2025(2): 35-39.

Tirana University
Faculty of Natural Sciences
Department of Biology

Integrated bibliography of publications and online resources on Albanian Lepidoptera.

Publication date: 31.viii.2025.
DOI: https://doi.org/10.5281/zenodo.15761299

Cuvelier Sylvain¹ | Paparisto Anila²

^{1^*}Diamantstraat 4, B-8900 Ieper, Belgium.
² Faculty of Natural Sciences, Department of Biology, Bulevardi Zogu i Pare, Tirana University, Al-1001 Tiranë, Albania.
^* Corresponding author: sylvain.cuvelier@telenet.be

Abstract
Albania’s butterfly fauna has been increasingly documented over the past century, with research evolving from early descriptive studies to comprehensive modern investigations.
The Fluturat e Shqipërisë website consolidates this growing body of work, integrating scientific publications, checklists, and monitoring data into a single resource.
In this publication, the referenced sources are systematically organized chronologically and alphabetically, reflecting the multidisciplinary and international scope of Lepidoptera research relevant to Albania and supporting ongoing scientific and conservation efforts.

Key words
Albania, Lepidoptera, data compilation, bibliography, online resources, Fluturat e Shqipërisë.

Introduction
Research on the butterfly fauna of Albania began in the early 20th century, with the earliest published contributions primarily authored by Prof. H. Rebel
A more sustained scientific interest developed after 1990, notably through the work of the late Prof. K. Misja, as well as contributions by S. Abadjiev and S. Beshkov, who laid important groundwork for the documentation of Albania’s Lepidoptera.
In recent years, the documentation of butterfly diversity in Albania has increased substantially, resulting in numerous publications and data sources covering taxonomy, distribution, ecology, and conservation. The website Fluturat e Shqipërisë compiles and integrates this growing body of literature, providing a centralized resource based onscientific articles, faunal checklists and biodiversity monitoring platforms.
The following list includes all sources referenced in the development of Fluturat e Shqipërisë, arranged chronologically and alphabetically by author or institutional source. It reflects the multidisciplinary and international scope of current Lepidoptera research relevant to the Albanian fauna.

Sources

2025	Cuvelier S. & Marafi M. AJ. 2025. Status and conservation of genetic diversity in the disjunct populations of Pseudochazara tisiphone Brown, [1981], in Albania. — Archives of Western Palearctic Lepidoptera 2025(1): 71-78 (pdf) (url)
	Cuvelier S., Marafi M. AJ. & Paparisto A. 2025. Synonymy of Pseudochazara misjai and Pseudochazara tisiphone dibra: taxonomic reassessment of an Albanian butterfly population. Is the glass half full or half empty? An inconvenient truth. — Archives of Western Palearctic Lepidoptera 2025(2): 71-20. 10.5281/zenodo.15492634 (url)
	Jakšić P. 2025. The Butterflies and Skippers of the Kosovo and Metohija, Republic of Serbia А Faunal and Ecological study. — Institute for Biological Research "Siniša Stanković" – National Institute of Republic of Serbia, University of Belgrade, 277 p. (url)
	Nahirnić-Beshkova A. 2025. An updated checklist of Psychidae (Lepidoptera) of Albania. — Historia naturalis bulgarica 47(5) 101-106. https://doi.org/10.48027/hnb.47.052
	Parmentier L. & Qirinxhi X.* 2025. Unexpected cryptic diversity revealed through integrative analysis within isolated populations of the Graylings (Lepidoptera: Nymphalidae: Satyrinae) in the Western Balkans — Phegea 53(1) 22-38. (accessed 01.iii.2025, abstract) *Qirinxhi is listed as a co-author on the publication, yet she has confirmed that she had no involvement in the article based on ICMJE and COPE authorships criteria.
	Parmentier L., Qirinxhi X. * & De Prins W. 2025. Precedence between simultaneously published new taxa names in the genus Pseudochazara (Nymphalidae: Satyrinae) with a synonymisation act based on the rules of the International Code of Zoological Nomenclature. — Phegea 53(2): 93-96. (accessed via Figshare on 02.viiii.2025, version 1) *Qirinxhi is listed as a co-author on the publication, yet she has once again confirmed that she had no involvement in the article based on ICMJE and COPE authorships criteria.
	Taymans M. & Cuvelier S. 2025. A dynamic checklist of the Western Palearctic butterflies, hyperlinked to the original descriptions at species, genus and family level (Lepidoptera, Papilionoidea). — Archives of Western Palearctic Lepidoptera 2025(1): 1-70 (pdf) (url)
	Verhulst J. 2025. Un périple de 20 jours dans les Balkans, du 26 juin au 14 juillet 2024. — Lambillionea CXXV (1) 18-35. (url)

2024	Clarke H. 2024. A checklist of European butterfly larval foodplants. — Ecology and evolution 14, e10834. (url)
	Miho A., Bego F. & Beqiraj S. 2024. Vlerat natyrore dhe biodiversiteti i Deltës së Vjosës – gjendja sot dhe kërcënimet. Natural values and biodiversity of the Vjosa Delta - current situation and threats. Përmbledhje punimesh / Collected works. FShN, UT, Tiranë / FNS, UT, Tirana (url)
	Peqini L., Paparisto A., Elezaj E., Tatari F., Cuvelier S. & Miho A. 2024 in Miho A., Bego F. & Beqiraj S. 2024. Vështrim mbi insektet (Klasa Insecta) e Deltës së Vjosës / Overview on the insects (Class Insecta) of the Vjosa Delta (url)
	Qirinxhi X., Halimi E., Paparisto A., Elena H. & Zaharia B.. 2024.A Perspective in the Study of Biodiversity of Order Lepidoptera in Albania: Case Study using the MRR Methodology. — International Journal of Advanced Multidiscipllinary Research and Studies 4(2): 1533-1536 (url)
	Striniqi A. 2024. Assessment of the endangered Lepidopterafauna of the Shkodra Lake area as a factor in preserving the biodiversity values of this ecosystem. — International Journal of Ecosystems and Ecology Science (IJEES) 14(4): 121-128 (url)
	Taymans M. & Cuvelier S. 2024. A dynamic checklist of the Western Palearctic butterflies, hyperlinked to the original descriptions at species, genus and family level (Lepidoptera, Papilionoidea). — Lépidoptères 33(84): 18-21 (url)

2023	Barkmann F., Huemer P., Tappeiner U., Tasser E. & Rüdisser J. 2023. Standardized butterfly surveys: comparing transect counts and area-time counts in insect monitoring. — Biodiversity and Conservation. https://doi.org/10.1007/s10531-022-02534-2 (url)
	Blicher Bjerregård E., Bjerg M., Clausen A., Kaj K., Glintborg S. & Videnkjær C. 2023. Butterflies of Albania: new surveys, four new records and a new checklist (Lepidoptera: Papilionoidea). — Phegea 51(2): 65–76. 10.6084/m9.figshare.22014704
	Bond K. 2023. Lepidoptera recorded in Albania during two visits, 2009 and 2014. — Entomologist’s Record and Journal of Variation 135(3): 129–136.
	Cuvelier S. 2023. Albania, a country with unexpected, intraspecific genetic variability in butterflies (Papilionoidea: Nymphalidae & Lycaenidae). Balancing on a tightrope between species, subspecies, ESU's and haplotypes. — Lépidoptères 32(82): 32-40 (url)
	Cuvelier S., Parmentier L., Qirinxhi X. & Paparisto A. 2023. Butterflies of Albania new data and going online. Fluturatat e Shqipërisë të dhëna të reja dhe faqja online (Lepidoptera: Papilionoidea) — Buletini i Shkencave te Natyres, Tirana university. 32(2022): 5-31 (url)
	Engelhardt E., Bowler D. & Hof C. 2023. European Habitat Directive has fostered monitoring but not prevented species declines. — Conservation Letters 16 e12948. (url)
	Kawahara A. et al. 2023. A global phylogeny of butterflies reveals their evolutionary history, ancestral hosts and biogeographic origins. — Nature ecology and evolution 7: 903-913. (url)
	Malatesta L., De Sanctis M., Ammann E., Attore F., Buffi F., Cambria V., Fratarcangeli C., Hoda P., Mahmutaj E., Meco M., Mullaj A., Shuka L. & Fanelli G. 2023. Bioregionalization of Albania: Mismatch between the flora and the climate suggests that our models of Southern European bioregions are in need of a revision. Folia Geobotanica (url)
	Prendi M., Paparisto A. & Cuvelier S. 2023. Proterebia phegea (Lepidoptera: Nymphalidae: Satyrinae): building bridges between the relic populations of Croatia, Bosnia-Herzegovina, Greece, and a new record from northern Albania. — Phegea 51(2): 59–64 (url)

2022	Clarke H. 2022. A provisional checklist of European butterfly larval foodplant. — Nota Lepidopterologica 45: 139-167 (url)
	Dapporto L., Menchetti M., Vodă R., Corbella C., Cuvelier S., Djemadi I., Gascoigne-Pees M., Hinojosa J., Lam N., Serracanta M., Talavera G., Dincă V. & Vila. R. 2022a. The Atlas of mitochondrial diversity of Western Palearctic butterflies. — Global Ecology and Biogeography. 00, 1–7 (url)
	Dapporto L., Menchetti M., Vodă R., Corbella C., Cuvelier S., Djemadi I., Gascoigne-Pees M., Hinojosa J., Lam N., Serracanta M., Talavera G., Dincă V. & Vila. R. 2022b. The Atlas of mitochondrial diversity of Western Palearctic butterflies. Appendix S1. — Global Ecology and Biogeography. 00, 1–7 (url)
	eBMS - mobile application. (url) (consulted 20.ii.2022)
	European Butterfly Monitoring Scheme (eBMS) website. (url) (consulted 20.ii.2022)
	Kabashi-Kastrati E., Zhushi-Etemi F., Çadraku H. & Bytyçi P. 2022. First record of Heteropterus morpheus (Pallas, 1771) (Lepidoptera: Hesperiidae) from the Republic of Kosovo. – Ecologia Balkanica 14(1): 149-153 (url)
	Kabashi-Kastrati E., Zhushi-Etemi F. & Bytyçi P. 2022. Notes on the presence of Erebia alberganus (De Prunner, 1879) (Lepidoptera: Nymphaliddae) for Kosovo. – Polish Journal of Natural Sciences 37(4): 445-453 (url)
	Plebejus orphicus (Kolev, 2005) in The Butterflies of Bulgaria. (url)
	Miljević, M., Tot I. & Djurić M. (2014-2022) Alciphron - database on insects of Serbia (butterflies) (url) (consulted 20.ii.2022)
	Nahirnić-Beshkova A., Beshkov S. & Jakšić P. 2022. New records of rare butterflies (Lepidoptera: Papilionoidea) in Albania. — Historia naturalis bulgarica 44(10): 109–115 (url)
	NC STATE Agriculture and Life Sciences. (url) (consulted 12.viii.2022)
	Pamperis L. 2022. New Maps WGS84 of distribution (url) (consulted 12.viii.2022)
	Parmentier L. 2022. Contribution to the Lepidopteran fauna of Albania with a first confirmation of Satyrium pruni (Linnaeus, 1758) (Lepidoptera: Lycaenidae) present in the country. — Phegea 50(3): 160-163 (url)
	Parmentier L., Vila R. & Lukhtanov V. 2022. Integrative analysis reveals cryptic speciation linked to habitat differentiation within Albanian populations of the anomalous blues (Lepidoptera, Lycaenidae, Polyommatus Latreille, 1804) – Comparative Cytogenetics 16(4): 211-242.2022 (url)
	Raymond B. & Gibbons R. Identification Guide to the Pyrgus Group of Grizzled Skippers in Europe (© Copyright Butterfly Conservation) (url) (consulted 12.viii.2022)
	Verovnik R. & Verovnik J. 2022. First record of Proterebia phegea (Lepidoptera: Satyrinae) from Albania. — Natura Croatica 31(1): 115–120 (url)

2021	Beshkov S. & Nahirnić-Beshkova. 2021. Contribution to knowledge of the Balkan Lepidoptera II (Lepidoptera: Macrolepidoptera) — Ecologica Montenegrina 42: 1–44 (url)
	Dincă V., Dapporto L., Somervuo P., Vodă R., Cuvelier S., Gascoigne-Pees M., Huemer P., Mutanen M., Hebert P. & Vila R. 2021. High resolution DNA barcode library for European butterflies reveals continental patterns of mitochondrial genetic diversity. — Communications Biology (url)
	Koren T., Kulijer D., Vukotić K. & Zhushi-Etemi F. 2021. Contribution to the knowledge of the butterfly diversity (Lepidoptera: Papilionoidea) of Kosovo. — Entomologist’s Gazette 72: 119-129 (url)
	Mølgaard M. & Cuvelier S. 2021. Web application for recognition of Greek butterflies - a review. — Nordjysk Lepidopterolklub 40(3): 9-17 (pdf)
	Warren et al. 2021. The decline fo butterflies in Europe: Problems, significance, and possible solutions. — PNAS 118(2) e2002551117. https://doi.org/10.1073/pnas.2002551117 (url)

2020	Bozhinovska E. & Melovski D. 2020. Distribution of butterflies Lepidoptera, Papilionidae & Hesperoidea in the Prespa region, North Macedonia, Albania and Greece. — Bulletin of the Biology Student’s Research Society 5: 97-110 (url)
	Hinojosa J. Dapporto L., Brockmann E., Dinca V., Tikhonov V., Grishin N., Lukthanov V. & Vila R. 2020. Overlooked cryptic diversity in Muschampia (Lepidoptera: Hesperiidae) adds two species to the European butterfly fauna. — Zoological Journal of the Linnean Society. 193(3): 847-859. (url)
	Shuka D., Tan K., Hallaçi B. & Shuka L. 2020. Additions to the flora of North Albania. — Phytologia Balcanica, 26(3): 517–522 (url)

2019	Aistleitner E. 2019. Coenonympha-Studien 2: Anmerkungen zu Arten der Gattung Coenonympha Hubner, 1819 (Lepidoptera: Nymphalidae: Satyrinae) – Nachrichten des Entomologischen Vereins Apollo. 40(2): 75–85.
	Kudrna O. 2019. Distribution of Butterflies and Skippers in Europe (Lepidoptera: Rhopalocera, Grypocera). 24 years Mapping European Butterflies (1995-2019). Final Report. Společnost pro Ochranu Motýlů, Prachatice, Czech Republic, 363 p.
	Maes et al. 2019. Integrating national Red Lists for prioritising conservation actions for European butterflies. — Journal of Insect conservation. https://doi.org/10.1007/s10841-019-00127-z (url)
	Striniqi-Laçej A. & Misja K. 2019. A review of threatened and risked entomophauna on Shkodra Lake area. —International Journal of Ecosystems and Ecology Science 9(2): 257–264 (url)
	van Swaay C. et al. 2019. The EU butterfly indicator for grassland species: 1990-2017: technical report. Butterfly Conservation Europe & ABLE/eBMS (www.butterfly-monitoring.net) (url)

2018	Barina Z., Somogyi G., Pifko D. & Rakaj M. 2018. Checklist of vascular plants of Albania. — Phytotaxa 378(1): 001-339 (url)
	Cuvelier S., Parmentier L., Paparisto A. & Couckuyt J. 2018. Butterflies of Albania – Fluturat e Shqipërisë. New surveys, new species and a new checklist (Lepidoptera: Papilionoidea) — Phegea 46(2): 48-69 (url)
	Cuvelier S., Parmentier L., Paparisto A. & Couckuyt J. 2018. Butterflies of Albania – Fluturat e Shqipërisë. New surveys, new species and a new checklist (Lepidoptera: Papilionoidea) S4: distribution maps for all species. — Phegea 46(2) S4 (url)
	Franeta F. 2018. Checklist of the butterflies (Lepidoptera: Papilionoidea) of Montenegro. — Zootaxa 4392: 128-148 (url)
	van Eldijk T. Wappler T., Strother P., van der Weijst C., Rajaei H., Visscher H. & van de Schootbrugge. 2018. A Triassic-Jurassic window into the evolution of Lepidoptera. — Science Advances 4(1), e1701568. DOI: 10.1126/sciadv.1701568 (url)
	Vrenozi B. & Aistleitner E. 2018. Distribution and biotope preference of Melanargia galathea (Linnaeus, 1758) in Albania (Lepidoptera, Nymphalidae, Satyrinae) – Buletini i Shkencave te Natyres (Universiteti i Tiranes) 25: 31-38 (url)
	Wiemers M., Balletto E., Dincă V., Fric Z., Lamas G., Lukhtanov V., Munguira M., van Swaay C., Vila R., Vliegenthart A., Wahlberg N. & Verovnik R. 2018. An updated checklist of the European Butterflies (Lepidoptera, Papilionoidea) — ZooKeys 811: 9-45 (url)

2017	de Jong R. & Coutsis J. 2017. A re-appraisal of Gegenes Hübner, 1819 (Lepidoptera, Hesperiidae) based on male and female genitalia, with the description of a new genus, Afrogegeness. — Tijdschrift voor Entomologie 160: 41-60 (url)
	Etemi F., Bytyqi P. & Visoka V. 2017. New Records of Araschnia levana (Linneaus, 1758) (Lepidoptera: Nymphalidae) in Kosovo. — Journal of International Environmental Application & Science 12(1): 5–8 (url)
	Hallman C., Sorg M., Jongejans E., Siepel H., Hofland N., Schwan H. et al. 2017. More than 75 percent decline over 27 years in total flying insect biomass in protected areas. — Plos One 12(10) e0185809. https://doi.org/10.1371/journal.pone.0185809 (url)
	Cuvelier S. & Maertens D. 2017. Trial test of external morphology-based identification of Leptidea sinapis, L. reali and L. juvernica (Lepidoptera: Pieridae) provides opportunity for an online identification platform. — Phegea 45(2): 41-45. (url)
	Mitter C., Davis D. & Cummings M. 2017. Phylogeny and Evolution of Lepidoptera. — Annual Review of Entomology 62: 265-283 (url)
	Nahirnić-Beshkova A. & Beshkov A. 2017. Penestoglossa dardoinella (Millière, 1863) (Lepidoptera: Psychidae) recorded for the first time in Albania. — Zoonotes 101: 1-3 (url)
	ASIG Geoportal. — https://geoportal.asig.gov.al/ (last consulted on 24.vi.2017).

2016	Cuvelier S., De Mol R. & Mølgaard M. 2016. Two extreme forms of Pseudochazara amymone (Lepidoptera: Nymphalidae, Satyrinae) from Albania. — Phegea 44(1): 14–16 (url)
	Dhora Dh., Dhora D. & Dhora A. 2016. Liqeni i Shkodrës. — Shtëpia Botuese Fiorentia, Shkodër. 202 p.
	Kolev Z. & Shtinkov N. 2016. The Pygmy Skipper Gegenes pumilio: a new species to Bulgaria, and a confirmation of its occurrence in the eastern Balkan Peninsula (Lepidoptera: Hesperiidae). – Phegea 44(1): 17-22 (url)
	Numa C., van Swaay C., Wynhoff I., Wiemers M., Barrios V., Allen D., Sayer C., López Munguira M., Balletto E., Benyamini D., Beshkov S., Bonelli S., Caruana R., Dapporto L., Franeta F., Garcia-Pereira Karaçetin E., Katbeh-Bader A., Maes D., Micevski N., Miller R., Monteiro E., Moulai R., Nieto A., Pamperis L., Pe’er G., Power A., Šašić M., Thompson K., Tzirkalli E., Verovnik R., Warren M. & Welch H. 2016. The status and distribution of Mediterranean butterflies. — IUCN, Malaga, Spain. 32 p (url)
	Russell P. & Pateman J. 2016. Further observations on Melitaea ornata Christoph, 1893 (Lepidoptera: Nymphalidae) in the Republic of Macedonia. — Entomologist’s Gazette 67: 15–22 (url)
	Sachanowicz K., Luczkowski S. & Larysz A. 2016. State of Knowledge of Butterfly Fauna of Albania, with Three New Species for the Country. — Acta zoologica bulgarica 68(4): 511–518 (url)
	Shtinkov N., Kolev Z., Vila R. & Dincă V. 2016. The sibling species Leptidea juvernica and L. sinapis (Lepidoptera, Pieridae) in the Balkan Peninsula: ecology, genetic structure, and morphological variation. — Zoology 119: 11–20 (url)
	Sobczyk R. & Gligorović B. 2016. Diversity of Butterflies in the Zeta-Skadar Plain, Montenegro. — Acta Zoologica Bulgarica 68(2): 183– 190 (url)
	Švara V., Popović M., Peternel A., Radevski Đ., Vukotić K. & Verovnik R. 2016. Surveys of butterfly and skipper fauna in the southwestern part of the Republic of Macedonia (Lepidoptera: Papilionoidea & Hesperioidea). — Natura Sloveniae 18(2): 23–37 (url)
	Takáts K. & Mølgaard M. 2016. Partial mtCOI-sequences of Balkanic species of Pseudochazara (Lepidoptera: Nymphalidae, Satyrinae) reveal three well-differentiated lineages. — Entomologica romanica 19: 21–40 (url)
	Verovnik R. & Wiemers M. 2016. Species delimitation in the Grayling genus Pseudochazara (Lepidoptera, Nymphalidae, Satyrinae) supported by DNA barcodes. — Zookeys 600: 131–154 (url)

2015	Bink F. & Moenen R. 2015. Butterflies in the Benelux. Based on Dagvlinders in de Benelux. Revised and extended (url)
	Cuvelier S. & Mølgaard M. 2015. Pseudochazara amymone (Lepidoptera, Nymphalidae) in Albania: Variability analysis, androconial scales and new distributional data. — Nota lepidopterologica 38(1): 1–22 (url)
	Kudrna O., Pennerstorfer J. & Lux K. 2015. Distribution Atlas of European butterflies and skippers. — Wissenschaftlicher Verlag Peks i.K., Schwanfeld, Germany, 632 p.
	Micevski N. 2015. De Prunner’s Ringlet Erebia triaria (de Prunner, 1798) (Lepidoptera : Nymphalidae, Satyrinae) – a new species for the Republic of Macedonia. — Natura Sloveniae 16(1): 53–58 (url)
	Micevski N., Franeta F., Gascoigne-Pees M., Micevski B. & Verovnik R. 2015. Butterfly surveys in Albania during 2014 including the discovery of two new species for the country. — Ecologica Montenegrina 3: 1–12 (url)
	Nahirnić A., Jakšić P. & Viborg A. 2015. Colias caucasica balcanica (Pieridae) rediscovered in Montenegro, with additional records for Serbia. — Phegea 43(1): 6–10 (url)
	Russell P., Zitnan D. & Major V. 2015. Confirmation of the presence of Melitaea ornata Christoph, 1893 (Lepidoptera: Nymphalidae) in Macedonia (FYROM) and its host-plants 2015. — Entomologist’s Gazette 66: 13–24.
	Russell, P. 2015. Melitaea ornata Christoph, 1893 (Lepidoptera: Nymphalidae) discovered in Montenegro, with a list of butterflies recorded there in May 2013. — Entomologist’s Gazette 66: 199–204 (url)
	Šašić M., Popović M., Cuvelier S., Đurić M., Franeta F., Gascoigne-Pees M., Koren T., Maes D., Micevski B., Micevski N., Mølgaard M., van Swaay C., Wynhoff I. & Verovnik R. 2015. Contribution to the knowledge of the butterfly fauna of Albania. — Nota lepidopterologica 38(1): 29–45 (url)
	Shtinkov N., Kolev Z., Vila R. & Dincă V. 2015. The sibling species Leptidea juvernica and L. sinapis (Lepidoptera, Pieridae) in the Balkan Peninsula: ecology, genetic structure, and morphological variation. — Zoology. (url1, url2)
	Švara V., Zaksek B. & Verovnik R. 2015. Contribution to the knowledge of the butterfly fauna of Montenegro (Lepidoptera: Rhopalocera). — Acta Entomologica Slovenica 23(1): 37–48 (url)

2014	Abdija X. 2014. Vlerësimi sistematiko-ekologjik i ropalocerofaunës lerësimi të Malit Sharr dhe Rrethinave të tij në rrafshin e entomofaunë Maqedonase. — Doktoratura Universitetii Tiranës, fakulteti i shkencave natyrore. 144 pp. [in Albanese].
	Cuvelier S. 2014. The Butterflies and Skippers of the North-Aegean Islands of Híos, Psará and Lésvos (Greece). New observations and an update on their distribution (Papilionoidea & Hesperioidea). — Phegea 42(3): 58-70 (url)
	Gascoigne-Pees M., Verovnik R., Franeta F. & Popović M. 2014. The lifecycle and ecology of Pseudochazara amymone (Brown, 1976) (Lepidoptera: Nymphalidae, Satyrinae). — Nachrichten des Entomologischen Vereins Apollo, N. F. 35(3): 129–138 (url)
	Jugovic J. & Koren T. 2014. Wing pattern morphology of three closely related Melitaea (Lepidoptera, Nymphalidae) species reveals highly inaccurate external morphology-based species identification. — Nota Lepidopterologica 37(1): 75-90. (url)
	Louy D., Habel J., Ulrich W. & Schmitt T. 2014. Out of the Alps: The biogeograpy of a disjunctly distributed mountain butterfly, the Almond eyed ringlet Erebia alberganus (Lepidoptera, Satyrinae) — Journal of Heredity 105: 28–38 (url)
	Melovski D. 2014. Diversity of butterflies (Lepidoptera: Papilionoidea & Hesperioidea) in Prespa National Park (Albania) — Macedonian Ecological Society report, 15 p (url) (consulted 08.viii.2022)
	Melovski D. & Bozhinovsk E. 2014. New records for four butterfly species (Lepidoptera: Papilionoidea & Hesperioidea) in the Republic of Macedonia. — Journal of Natural Sciences Research 4(7): 40–44 (url)
	Verovnik R., Popovic M., Šašić M., Cuvelier S. & Maes D. 2014. Wanted! Dead or alive: the tale of the Brown’s Grayling (Pseudochazara amymone). — Journal of Insect Conservation 18(4): 675–682 (url)
	Zeneli G., Shuka L., Bego F., Kashta L., Shumka S., Hoxha S., Diku A., Bano E. & Krasniqi F. 2014. National Biodiversity Strategy of Albania (2012–2020). — Tirana, 146 p. (in English) (url)

2013	Abdija X., Beadini N., Beadini S. & Iseni A. 2013a. Preliminary data for the familia Nymphalidae of Sharr Mountain and its surroudings (Mavrovo and Pollog). — Anglisticum Journal (IJLLIS) 2(2): 227–236 (url)
	Abdija X., Beadini N. & Beadini S. 2013b. The basic characteristics of the study of butterfly (Rhopalocera) fauna in the mountain massif of Sharr, Pollog valley and the National Park of Mavrovo. — Advances in Life Science and Technology 8: 28–34 (url)
	Abdija X., Beadini N., Beadini S. & Rexhepi B. 2013c. A taxonomic study of the family Hesperiidae (Rhopalocera) of the massif mountains of Sharr, Pollog Valley and Mavrovo National Park. — Journal of Natural Sciences Research 3(12): 47–51 (url)
	Aistleitner E. 2013. Zur Kenntnis der Schmetterlingsfauna der Südalpen und der angrenzenden Dinariden (2) Der Mohrenfalter Erebia medusa (Denis & Schiffermüller, 1775), Taxonomie und Chorologie sowie Beschreibung einer neuen Unterart (Lepidoptera, Satyridae) – Atalanta 44(1/2): 119-125 (url)
	Bink F. & Moenen R. 2013. Dagvlinders in de Benelux. — Bink, Bennekom, 191 p.
	Hubrechts S. 2013. A re-evaluation of the cryptic species complex Leptidea (Lepidoptera, Pieridae) in Belgium. — Universiteit Antwerpen, Faculteit Wetenschappen, Evolutionaire ecologie, Antwerpen, 53 p.
	John E., Wiemers M., Makris C. & Russell P. 2013. The Pontia daplidice (Linnaeus, 1758)/Pontia edusa (Fabricius, 1777) complex (Lepidoptera: Pieridae): confirmation of the presence of Pontia daplidice in Cyprus, and of Cleome iberica DC. as a new host-plant for this species in the Levant. — Entomologist’s Gazette 64: 69–78 (url)
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2009	Cuvelier S. 2009. Pelopidas thrax, a new species for the Island of Kós and an update of its distribution in Greece (Lepidoptera: Hesperioidea & Papilionoidea). — Phegea 37(3): 84-93 (url)
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2008	Dincă V., Szekely L., Kovács S., Kovács Z. & Vila R. 2008. Pyrgus andromedae (Wallengren, 1853) (Hesperiidae) in the Romanian Carpathians. — Nota lepidopterologica 31(2): 263–272 (url)
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2007	Jakšić P. 2007. Contribution to knowledge of the butterflies of Mt. Paštrik, Serbia (Lepidoptera: Hesperioidea and Papilionoidea). — Acta entomologica serbica 12(2): 55–61 (url)

2006	Micevski N. & Micevski B. 2006. A biogeographical and ecological analysis of the Butterfly fauna of Pelister National Park. — Biologia Macedonica 57/58: 89–98 (url)
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2005	Coutsis J. & De Prins J. 2005. A new brown Polyommatus (Agrodiaetus) from northern Greece (Lepidoptera: Lycaenidae) — Phegea 33(4): 129-137. (url)
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1998	Jakšić P. 1998. Dnevni leptiri (Lepidoptera: Hesperioidea i Papilionoidea) Šar-planine. The butterflies of Šar-planina Mt. (Lepidoptera: Hesperioidea & Papilionoidea). — Zaštita prirode 50: 229–252 (url)

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Conclusion
This bibliography aims to offer the most complete overview to date of published and digital resources concerning the butterflies of Albania.
It acknowledges the valuable contributions of all researchers whose work has advanced the understanding of the country’s Lepidoptera.
To ensure ongoing accuracy and completeness, Fluturat e Shqipërisë remains open to updates and welcomes additional references that may have been overlooked or are newly published.

Author contributions
Both authors were equally involved in all stages of the research process, from conceptualization through data analysis, to drafting and revising the manuscript.

Acknowledgements
We acknowledge the foundational work of early researchers who initiated the study of Albania’s Lepidoptera. Their initial documentation provided a basis for continued scientific investigation. We also recognize the sustained efforts of subsequent authors and researchers whose publications have significantly contributed to expanding knowledge on the biodiversity of butterflies in Albania. Their collective work has been essential in building the current comprehensive understanding of the region’s Lepidoptera fauna and continue to inspire ongoing research and conservation in Albania’s rich natural heritage.

Chronicles Fluturat e Sqhipërisë 2025(2): 35-39.
https://doi.org/10.5281/zenodo.15761299
An open Access Article, which permits use, distribution and reproduction in any medium, provided the original work is properly cited.
© 2025. The Authors.